A.T. Robertson Commentary Revelation 4:9

"And when the living creatures shall give glory and honor and thanks to him that sitteth on the throne, to him that liveth for ever and ever," — Revelation 4:9 (ASV)

When the living creatures shall give (οταν δωσουσιν τα ζωα). Indefinite temporal clause with οταν and the future active indicative (δωσουσιν) rather than the more common second aorist active subjunctive (δωσιν) with the notion of repetition rather than unbroken continuance, "whenever they give." The giving of praise and glory to God by the four living creatures (representatives of nature) is met by corresponding worship by the redeemed (the four and twenty elders). "Created life adores the Uncreated" (Swete), "to the one living for ages of ages."